CLONING, EXPRESSION, AND CHARACTERIZATION OF A NOVEL MOLLUSK-DERIVED GROWTH FACTOR IN APLYSIA

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dc.contributor Corey S . Goodman
dc.contributor Sherry D . Painter
dc.contributor Darrell H . Carney
dc.contributor Jane E . Bottenstein
dc.contributor Gregg T . Nagle
dc.creator Akalal , David -Benjamin Garcia
dc.date 2003 -03 -27
dc.date.accessioned 2011 -04 -27T21 :36 :29Z
dc.date.available 2011 -04 -27T21 :36 :29Z
dc.date.issued 2011 -04 -27T21 :36 :29Z
dc.identifier http : / /etd .utmb .edu /theses /available /etd -01162003 -093008 /
dc.identifier.uri http : / /hdl .handle .net /2249 .1 /35653
dc.description While the mollusk Aplysia californica has been used as an excellent model for studying the nervous system , no known growth and /or neurotrophic factors had been discovered or characterized in this model . The discovery of an Aplysia growth factor would be important for studying the mechanisms of actions of growth factors and their roles in growth , development , and metamorphosis . In 1996 , an insect -derived growth factor (IDGF ) was purified from the conditioned medium of an embryonic fleshfly cell line . It had no significant homology to any other protein except for an Aplysia protein of unknown function , atrial gland granule -specific antigen (AGSA ) . Based on its similarity to IDGF , this project was aimed at determining if AGSA functions as a growth factor in Aplysia . In this study , we cloned an atrial gland cDNA , corrected the reported AGSA nucleotide sequence , and renamed the predicted protein mollusk -derived growth factor (MDGF ) . We have confirmed that there is significant MDGF gene expression in the Aplysia atrial gland and no detectable expression in acutely dissected adult CNS . However , we demonstrate that expression of the MDGF gene begins during embryonic development , and displays distinct temporal patterns of expression in the reproductive tract and CNS during juvenile development suggesting that MDGF may be important during Aplysia development . In addition , MDGF is detected in axotomized adult CNS , suggesting a potential role for MDGF in injury repair . We affinity -purified native MDGF protein from Aplysia atrial glands and showed that it has potent adenosine deaminase (ADA ) and growth factor activity , inducing up to a two -fold increase in an insect cell proliferation assay . A number of MDGF -related proteins having 30 -50 % sequence identity among themselves have recently been reported including : a human protein involved in cat eye syndrome , several insect growth factors possessing adenosine deaminase activity , and proteins from several other organisms (pig , zebrafish , cow , and slime mold ) . Our findings support the idea that MDGF belongs to a new family of ADA -related growth factors and that it may be important during development and repair of the Aplysia CNS by regulating the levels of extracellular adenosine . MDGF is the first growth factor purified and characterized in Aplysia .
dc.format application /pdf
dc.language en
dc.publisher UTMB
dc.rights unrestricted
dc.rights I hereby certify that , if appropriate , I have obtained and attached hereto a written permission statement from the owner (s ) of each third party copyrighted matter to be included in my thesis , dissertation , or project report , allowing distribution as specified below . I certify that the version I submitted is the same as that approved by my advisory committee . I hereby grant to University of Texas Medical Branch or its agents the non -exclusive license to archive and make accessible , under the conditions specified below , my thesis , dissertation , or project report in whole or in part in all forms of media , now or hereafter known . I retain all other ownership rights to the copyright of the thesis , dissertation or project report . I also retain the right to use in future works (such as articles or books ) all or part of this thesis , dissertation , or project report .
dc.source http : / /etd .utmb .edu /theses /available /etd -01162003 -093008 /
dc.subject Anatomy and Neurosciences Cell Biology Program
dc.title CLONING , EXPRESSION , AND CHARACTERIZATION OF A NOVEL MOLLUSK -DERIVED GROWTH FACTOR IN APLYSIA
dc.type text

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CLONING, EXPRESSION, AND CHARACTERIZATION OF A NOVEL MOLLUSK-DERIVED GROWTH FACTOR IN APLYSIA. Available electronically from http : / /hdl .handle .net /2249 .1 /35653 .

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